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PRD Class
Families List  Genes List
The PRD class is the second largest of the homeobox gene classes in animal genomes; like the ANTP class these genes have only been found in animals. The PRD class derives its name from (but is not an abbreviation for) the Paired (Prd) gene of Drosophila. PRD class genes are defined phylogenetically - they form a robust monophyletic group in evolutionary trees of homeodomain sequences. The PRD homeobox class overlaps with the Pax gene class - some but not all Pax genes are in the PRD homeobox gene class, while the vast majority of PRD class homeobox genes are not Pax genes.

There are 9 Pax genes in the human or mouse genomes, defined by presence of a 128 amino acid Paired domain, and of these 7 are members of the PRD homeobox class by virtue of possessing a homeobox: complete homeobox for PAX3, 4, 6 and 7; partial homeobox for PAX2, 5 and 8 (the two human Pax genes lacking a homeobox entirely are PAX1 and PAX9; these are not included in this database). In addition to these 7 homeobox-containing Pax genes, there are a further 43 PRD class genes in the human genome. These 50 genes can be divided into 31 gene families. There are also 24 human pseudogenes generated from these genes, plus an unknown number of DUX repetitive sequences (Holland et al. 2007).

In amphioxus there are 5 Pax genes, and of these 3 are have a partial or complete homeobox (Pax-1/9 and Pon are the two lacking homeobox sequence and therefore excluded; Pon is present in Drosophila and amphioxus, but not human). In addition to these genes, there are a further 27 PRD class genes in amphioxus (Takatori et al. 2008).

Reference(s):
Holland PWH, Booth HAF, Bruford EA (2007) BMC Biology 5:47.
Takatori N et al (2008) Dev Genes Evol 218: 579-590.
 
Families List of Class PRD :
Alx(16)AprdA(1)AprdB(1)AprdC(1)AprdD(2)AprdE(1)Argfx(3)Arx(14)CG11294(3)
Dmbx(8)Dprx(8)Drgx(9)Dux(42)Esx(2)Gsc(14)Hbn(3)Hesx(5)Hopx(5)
Isx(5)Leutx(1)Mix(7)Nobox(5)Obox(36)Otp(12)Otx(24)Pax2/5/8(12)Pax3/7(24)
Pax4/6(24)Phox(17)Pitx(20)Prop(8)Prrx(14)Rax(14)Repo(4)Rhox(39)Sebox(3)
Shox(12)Tprx(5)Uncx(16)Vsx(18)      
 
Loci in PRD of All back to top..
Symbol/LocusNameFamilyOrganismRegulator
Alx3-lAlx3 likeAlxZebrafish 
alx4aaristaless-like homeobox 4aAlxZebrafish 
alx4baristaless-like homeobox 4bAlxZebrafish 
arxaristaless related homeoboxArxZebrafish 
bonbonnie and clydeMixZebrafish 
cart1cartilage paired-class homeoprotein 1AlxZebrafish 
chr04.1Alx likeAlxZebrafish 
chr05.1Gsc likeGscZebrafish 
crxcone-rod homeoboxOtxZebrafish 
Dmbx1-lDmbx1 likeDmbxZebrafish 
dmbx1adiencephalon/mesencephalon homeobox 1aDmbxZebrafish 
dmbx1bdiencephalon/mesencephalon homeobox 1bDmbxZebrafish 
drgxdorsal root ganglia homeoboxDrgxZebrafish 
gscgoosecoidGscZebrafish 
hesx1homeo box (expressed in ES cells) 1HesxZebrafish 
LOC100330956bonnie and clyde-likeMixZebrafish 
og9xOG9 homeobox geneMixZebrafish 
otpaorthopedia homolog aOtpZebrafish 
otpborthopedia homolog bOtpZebrafish 
otx1aorthodenticle homolog 1aOtxZebrafish 
otx1borthodenticle homolog 1bOtxZebrafish 
otx2orthodenticle homolog 2OtxZebrafish 
otx5orthodenticle homolog 5OtxZebrafish 
pax2bpaired box gene 2bPax2/5/8Zebrafish 
pax3apaired box gene 3aPax3/7Zebrafish 
pax3bpaired box gene 3bPax3/7Zebrafish 
pax4paired box 4Pax4/6Zebrafish 
Pax6-lPax6 likePax4/6Zebrafish 
pax6apaired box gene 6aPax4/6Zebrafish 
pax6bpaired box gene 6bPax4/6Zebrafish 
pax7apaired box gene 7aPax3/7Zebrafish 
pax7bpaired box gene 7bPax3/7Zebrafish 
Phox-lPhox likePhoxZebrafish 
phox2apaired-like homeobox 2aPhoxZebrafish 
phox2bpaired-like homeobox 2bPhoxZebrafish 
phox2b-lphox2b like PhoxZebrafish 
pitx1paired-like homeodomain transcription factor 1PitxZebrafish 
pitx2paired-like homeodomain transcription factor 2PitxZebrafish 
pitx3paired-like homeodomain transcription factor 3PitxZebrafish 
prop1PROP paired-like homeobox 1PropZebrafish 
prrx1apaired related homeobox 1aPrrxZebrafish 
prrx1bpaired related homeobox 1bPrrxZebrafish 
rx1retinal homeobox gene 1RaxZebrafish 
rx2retinal homeobox gene 2RaxZebrafish 
rx3retinal homeobox gene 3RaxZebrafish 
shoxshort stature homeoboxShoxZebrafish 
shox2short stature homeobox 2ShoxZebrafish 
uncxUNC homeoboxUncxZebrafish 
uncx4.1Unc4.1 homeoboxUncxZebrafish 
Uncxl-2Uncx like 2UncxZebrafish 
vsx1visual system homeobox 1 homolog, chx10-likeVsxZebrafish 
vsx2visual system homeobox 2VsxZebrafish 
wu:fi69e09wu:fi69e09NoboxZebrafish 
alr-1AristaLess (Drosophila homeodomain) Related 1ArxNematode 
ceh-10C. Elegans Homeobox 10VsxNematode 
ceh-17C. Elegans Homeobox 17PhoxNematode 
ceh-36C. Elegans Homeobox 36OtxNematode 
ceh-37C. Elegans Homeobox 37OtxNematode 
ceh-45C. Elegans Homeobox 45GscNematode 
ceh-53C. Elegans Homeobox 53Pax3/7Nematode 
ceh-8C. Elegans Homeobox 8RaxNematode 
dsc-1Defecation Suppressor of Clk-1VsxNematode 
eyg-1EYG (Drosophila eyegone) homolog 1Pax3/7Nematode 
pax-3PAX (Paired box) transcription factor 3Pax3/7Nematode 
T13C5.4 PhoxNematode 
ttx-1abnormal ThermoTaXis 1OtxNematode 
unc-30UNCoordinated 30PitxNematode 
unc-4UNCoordinated 4UncxNematode 
unc-42UNCoordinated 42PropNematode 
vab-3Variable ABnormal morphology 3Pax4/6Nematode 
Alx1 AlxMouse 
Alx3 AlxMouse 
Alx4 AlxMouse 
Arx ArxMouse 
Crx OtxMouse 
Dmbx1 DmbxMouse 
Dux DuxMouse 
Duxbl DuxMouse 
Esx1 EsxMouse 
Gm10391 DuxMouse 
Gm10394 DuxMouse 
Gm10678 OboxMouse 
Gm10679 OboxMouse 
Gm14543Rhox7cRhoxMouse 
Gm14544 RhoxMouse 
Gm3959 OboxMouse 
Gm3965 OboxMouse 
Gm3968 OboxMouse 
Gm3977 OboxMouse 
Gm3980 OboxMouse 
Gm3981 OboxMouse 
Gm3984 OboxMouse 
Gm3987 OboxMouse 
Gm3989 OboxMouse 
Gm3994 OboxMouse 
Gm3996 OboxMouse 
Gm3998 OboxMouse 
Gm4001 OboxMouse 
Gm4003 OboxMouse 
Gm4014 OboxMouse 
Gm4016 OboxMouse 
Gm4018 OboxMouse 
Gm4023 OboxMouse 
Gm4745 OboxMouse 
Gm4830 OboxMouse 
Gm4897 OboxMouse 
Gm4981 DuxMouse 
Gm5496 OboxMouse 
Gm5889 OboxMouse 
Gm6310Rhox7bRhoxMouse 
Gm7590Rhox7dRhoxMouse 
Gm8040 OboxMouse 
Gm8053 OboxMouse 
Gm8057 OboxMouse 
Gm8064 OboxMouse 
Gm8072 OboxMouse 
Gsc GscMouse 
Gsc2 GscMouse 
Hesx1 HesxMouse 
Hopx HopxMouse 
Isx IsxMouse 
LOC100417000 OboxMouse 
LOC100504180 DuxMouse 
Mixl1 MixMouse 
Nobox NoboxMouse 
Obox1 OboxMouse 
Obox2 OboxMouse 
Obox3 OboxMouse 
Obox5 OboxMouse 
Obox6 OboxMouse 
Otp OtpMouse 
Otx1 OtxMouse 
Otx2 OtxMouse 
Pax2 Pax2/5/8Mouse 
Pax3 Pax3/7Mousemmu-mir-1; mmu-mir-206
Pax4 Pax4/6Mouse 
Pax5 Pax2/5/8Mouse 
Pax6 Pax4/6Mouse 
Pax7 Pax3/7Mousemmu-mir-206; mmu-mir-486
Pax8 Pax2/5/8Mouse 
Phox2a PhoxMouse 
Phox2b PhoxMouse 
Pitx1 PitxMouse 
Pitx2 PitxMouse 
Pitx3 PitxMouse 
Prop1 PropMouse 
Prrx1 PrrxMouse 
Prrx2 PrrxMouse 
Prrxl1 DrgxMouse 
Rax RaxMouse 
Rhox1 RhoxMouse 
Rhox10 RhoxMouse 
Rhox11 RhoxMouse 
Rhox12 RhoxMouse 
Rhox13 RhoxMouse 
Rhox2a RhoxMouse 
Rhox2b RhoxMouse 
Rhox2c RhoxMouse 
Rhox2d RhoxMouse 
Rhox2e RhoxMouse 
Rhox2f RhoxMouse 
Rhox2g RhoxMouse 
Rhox2h RhoxMouse 
Rhox3-ps RhoxMouse 
Rhox3a RhoxMouse 
Rhox3b RhoxMouse 
Rhox3c RhoxMouse 
Rhox3e RhoxMouse 
Rhox3f RhoxMouse 
Rhox3g RhoxMouse 
Rhox3h RhoxMouse 
Rhox4a RhoxMouse 
Rhox4b RhoxMouse 
Rhox4c RhoxMouse 
Rhox4d RhoxMouse 
Rhox4e RhoxMouse 
Rhox4f RhoxMouse 
Rhox4g RhoxMouse 
Rhox6 RhoxMouse 
Rhox7 RhoxMouse 
Rhox8 RhoxMouse 
Rhox9 RhoxMouse 
Sebox SeboxMouse 
Shox2 ShoxMouse 
Uncx UncxMouse 
Vsx1 VsxMouse 
Vsx2 VsxMouse 
ALX1aristaless-like homeobox 1AlxHuman 
ALX3aristaless-like homeobox 3AlxHuman 
ALX4aristaless-like homeobox 4AlxHuman 
ARGFXarginine-fifty homeoboxArgfxHuman 
ARGFXP1arginine-fifty homeobox pseudogene 1ArgfxHuman 
ARGFXP2arginine-fifty homeobox pseudogene 2ArgfxHuman 
ARXaristaless related homeoboxArxHuman 
CRXcone-rod homeoboxOtxHuman 
DMBX1diencephalon/mesencephalon homeobox 1DmbxHuman 
DPRXdivergent-paired related homeoboxDprxHuman 
DPRXP1divergent-paired related homeobox pseudogene 1DprxHuman 
DPRXP2divergent-paired related homeobox pseudogene 2DprxHuman 
DPRXP3divergent-paired related homeobox pseudogene 3DprxHuman 
DPRXP4divergent-paired related homeobox pseudogene 4DprxHuman 
DPRXP5divergent-paired related homeobox pseudogene 5DprxHuman 
DPRXP6divergent-paired related homeobox pseudogene 6DprxHuman 
DPRXP7divergent-paired related homeobox pseudogene 7DprxHuman 
DRGXdorsal root ganglia homeoboxDrgxHuman 
DUX4double homeobox, 4DuxHuman 
Dux4l DuxHuman 
DUX4L10double homeobox 4 like 10DuxHuman 
DUX4L11double homeobox 4 like 11DuxHuman 
DUX4L12double homeobox 4 like 12DuxHuman 
DUX4L13double homeobox 4 like 13DuxHuman 
DUX4L14double homeobox 4 like 14DuxHuman 
DUX4L15double homeobox 4 like 15DuxHuman 
DUX4L16double homeobox 4 like 16DuxHuman 
DUX4L17double homeobox 4 like 17DuxHuman 
DUX4L18double homeobox 4 like 18DuxHuman 
DUX4L19double homeobox 4 like 19DuxHuman 
DUX4L2double homeobox 4 like 2DuxHuman 
DUX4L20double homeobox 4 like 20DuxHuman 
DUX4L3double homeobox 4 like 3DuxHuman 
DUX4L4double homeobox 4 like 4DuxHuman 
DUX4L5double homeobox 4 like 5DuxHuman 
DUX4L6double homeobox 4 like 6DuxHuman 
DUX4L7double homeobox 4 like 7DuxHuman 
DUX4L8double homeobox 4 like 8DuxHuman 
DUX4L9double homeobox 4 like 9DuxHuman 
DUXAdouble homeobox ADuxHuman 
DUXAP1double homeobox A pseudogene 1DuxHuman 
DUXAP10double homeobox A pseudogene 10DuxHuman 
DUXAP2double homeobox A pseudogene 2DuxHuman 
DUXAP3double homeobox A pseudogene 3DuxHuman 
DUXAP4double homeobox A pseudogene 4DuxHuman 
DUXAP5double homeobox A pseudogene 5DuxHuman 
DUXAP6double homeobox A pseudogene 6DuxHuman 
DUXAP7double homeobox A pseudogene 7DuxHuman 
DUXAP8double homeobox A pseudogene 8DuxHuman 
DUXAP9double homeobox A pseudogene 9DuxHuman 
DUXBdouble homeobox BDuxHuman 
DUXBLRDuxbl remnantsDuxHuman 
ESX1extraembryonic, spermatogenesis, homeobox 1 homolog (mouse)EsxHuman 
GSCgoosecoid homeoboxGscHuman 
GSC2goosecoid homeobox 2GscHuman 
HESX1HESX homeobox 1HesxHuman 
HOPXHOP homeoboxHopxHuman 
ISXintestine-specific homeoboxIsxHuman 
LEUTXleucine-twenty homeoboxLeutxHuman 
LOC100131369double homeobox, 4-like pseudogeneDuxHuman 
LOC100506764double homeobox protein 4-likeDuxHuman 
MIXL1Mix paired-like homeoboxMixHuman 
NOBOXNOBOX oogenesis homeoboxNoboxHuman 
OTPorthopedia homologOtpHuman 
OTX1orthodenticle homeobox 1OtxHuman 
OTX2orthodenticle homeobox 2OtxHuman 
OTX2P1orthodenticle homeobox 2 pseudogeneOtxHuman 
PAX2paired box gene 2Pax2/5/8Human 
PAX3paired box 3Pax3/7Human 
PAX4paired box gene 4Pax4/6Human 
PAX5paired box gene 5Pax2/5/8Human 
PAX6paired box 6Pax4/6Human 
PAX7paired box 7Pax3/7Human 
PAX8paired box gene 8Pax2/5/8Human 
PHOX2Apaired-like homeobox 2aPhoxHuman 
PHOX2Bpaired-like homeobox 2bPhoxHuman 
PITX1pituitary homeobox 1PitxHuman 
PITX2pituitary homeobox 2PitxHuman 
PITX3pituitary homeobox 3PitxHuman 
PROP1DROP paired-like homeobox 1PropHuman 
PRRX1paired related homeobox 1PrrxHuman 
PRRX2paired related homeobox 2PrrxHuman 
RAXretina and anterior neural fold homeoboxRaxHuman 
RAX2retina and anterior neural fold homeobox 2RaxHuman 
RHOXF1Rhox homeobox family, member 1RhoxHuman 
RHOXF2Rhox homeobox family, member 2RhoxHuman 
RHOXF2BRhox homeobox family, member 2BRhoxHuman 
SEBOXSEBOX homeoboxSeboxHuman 
SHOXshort stature homeoboxShoxHuman 
SHOX2short stature homeobox 2ShoxHuman 
TPRX1tetra-peptide repeat homeobox 1TprxHuman 
TPRX1P1tetra-peptide repeat homeobox 1 pseudogene 1TprxHuman 
TPRX1P2tetra-peptide repeat homeobox 1 pseudogene 2TprxHuman 
TPRX2Ptetra-peptide repeat homeobox 2 pseudogeneTprxHuman 
TPRXLtetra-peptide repeat homeobox-likeTprxHuman 
UNCXUNC homeoboxUncxHuman 
VSX1visual system homeobox 1VsxHuman 
VSX2visual system homeobox 2VsxHuman 
AlAristalessArxHoneybee 
CG11294CG11294CG11294Honeybee 
DrgxDrgxDrgxHoneybee 
EyEyelessPax4/6Honeybee 
EygEyegonePax4/6Honeybee 
GsbGooseberryPax3/7Honeybee 
GsbnGooseberry neuroPax3/7Honeybee 
GscGoosecoidGscHoneybee 
HbnHomeobrainHbnHoneybee 
Oc1Ocelliless1OtxHoneybee 
Oc2Ocelliless2OtxHoneybee 
OtpOrthopediaOtpHoneybee 
PhoxPhoxPhoxHoneybee 
Pph13Pph13ArxHoneybee 
PrdPairedPax3/7Honeybee 
PropPropPropHoneybee 
PrrxPrrxPrrxHoneybee 
Ptx1Ptx1PitxHoneybee 
RepoReversed polarityRepoHoneybee 
RxRetinal homeoboxRaxHoneybee 
ShoxShoxShoxHoneybee 
SvShavenPax2/5/8Honeybee 
ToyTwin of EyelessPax4/6Honeybee 
Unc4Unc4UncxHoneybee 
VsxVsxVsxHoneybee 
alaristalessArxFruitfly 
CG11294CG11294CG11294Fruitfly 
CG32532CG32532PropFruitfly 
CG34340CG34340DrgxFruitfly 
CG34367CG34367ShoxFruitfly 
CG9876CG9876PrrxFruitfly 
eyeyelessPax4/6Fruitfly 
eygeyegonePax4/6Fruitfly 
gsbgooseberryPax3/7Fruitfly 
gsb-ngooseberry-neuroPax3/7Fruitfly 
GscGoosecoidGscFruitfly 
hbnhomeobrainHbnFruitfly 
ococellilessOtxFruitfly 
OdsHOds-site homeoboxUncxFruitfly 
otporthopediaOtpFruitfly 
PHDPPutative homeodomain proteinPhoxFruitfly 
Pph13PvuII-PstI homology 13ArxFruitfly 
prdpairedPax3/7Fruitfly 
Ptx1Ptx1PitxFruitfly 
reporeversed polarityRepoFruitfly 
RxRetinal HomeoboxRaxFruitfly 
svshavenPax2/5/8Fruitfly 
toetwin of eygPax4/6Fruitfly 
toytwin of eyelessPax4/6Fruitfly 
tuptailupVsxFruitfly 
unc-4unc-4UncxFruitfly 
Vsx1 VsxFruitfly 
Vsx2 VsxFruitfly 
alx1ALX homeobox 1 AlxFrog 
alx4ALX homeobox 4 AlxFrog 
arxaristaless related homeobox ArxFrog 
arx-larx likeArxFrog 
crxcone-rod homeobox OtxFrog 
dmbx1diencephalon/mesencephalon homeobox 1 DmbxFrog 
drgxdorsal root ganglia homeoboxDrgxFrog 
gscgoosecoid homeoboxGscFrog 
hesx1HESX homeobox 1HesxFrog 
hopxHOP homeoboxHopxFrog 
isxintestine-specific homeoboxIsxFrog 
nobox-lnobox likeNoboxFrog 
otp-l OtpFrog 
otpa OtpFrog 
otpborthopedia homeobox OtpFrog 
otx1orthodenticle homeobox 1 OtxFrog 
otx2orthodenticle homeobox 2OtxFrog 
pax2 Pax2/5/8Frog 
pax4lpax4 homeobox likePax4/6Frog 
pax6paired box 6Pax4/6Frog 
pax6lpaired box 6 likePax4/6Frog 
pax7paired box 7Pax3/7Frog 
phox2apaired-like homeobox 2aPhoxFrog 
phox2bpaired-like homeobox 2b PhoxFrog 
pitx1paired-like homeodomain 1 PitxFrog 
pitx2paired-like homeodomain 2 PitxFrog 
pitx2b PitxFrog 
prrx1bpaired related homeobox 1PrrxFrog 
prrx2paired related homeobox 2 PrrxFrog 
raxretina and anterior neural fold homeobox RaxFrog 
rax2 retina and anterior neural fold homeobox 2 RaxFrog 
shoxshort stature homeobox ShoxFrog 
shox2short stature homeobox 2 ShoxFrog 
uncx1UNC homeobox 1UncxFrog 
uncx2UNC homeobox 2UncxFrog 
vsx1visual system homeobox 1VsxFrog 
vsx2visual system homeobox 2VsxFrog 
ALX1ALX homeobox 1AlxChicken 
ALX4ALX homeobox 4AlxChicken 
ARXaristaless related homeoboxArxChicken 
DMBX1diencephalon/mesencephalon homeobox 1DmbxChicken 
DRGXdorsal root ganglia homeoboxDrgxChicken 
GSCgoosecoid homeoboxGscChicken 
GSC2goosecoid homeobox 2GscChicken 
HESX1HESX homeobox 1HesxChicken 
HOPXHOP homeoboxHopxChicken 
ISXintestine-specific homeoboxIsxChicken 
LOC419032similar to newborn ovary homeodomain proteinNoboxChicken 
LOC769839Mix likeMixChicken 
LOC771564Otp likeOtpChicken 
MIXL1Mix1 homeobox-like 1MixChicken 
PAX3paired box 3Pax3/7Chicken 
PAX6paired box gene 6Pax4/6Chicken 
PAX7paired box 7Pax3/7Chicken 
PHOX2Bpaired-like homeobox 2bPhoxChicken 
PITX1paired-like homeodomain 1PitxChicken 
PITX2paired-like homeodomain 2PitxChicken 
PITX3paired-like homeodomain 3PitxChicken 
PRRX1paired related homeobox 1PrrxChicken 
PRRX2paired related homeobox 2PrrxChicken 
RX2retina and anterior neural fold homeoboxRaxChicken 
SEBOXSEBOX homeoboxSeboxChicken 
SHOX2short stature homeobox 2ShoxChicken 
UNCXUNC homeoboxUncxChicken 
VSX1visual system homeobox 1VsxChicken 
VSX2visual system homeobox 2VsxChicken 
AlAristalessArxBeetle 
CG11294CG11294CG11294Beetle 
DrgxDrgxDrgxBeetle 
EyEyelessPax4/6Beetle 
EygEyegonePax4/6Beetle 
GsbGsbPax3/7Beetle 
GsbnGooseberry neuroPax3/7Beetle 
GscGoosecoidGscBeetle 
HbnHomeobrainHbnBeetle 
Oc1Ocelliless1OtxBeetle 
Oc2Ocelliless2OtxBeetle 
OtpOrthopediaOtpBeetle 
PhoxPhoxPhoxBeetle 
Pph13PvuII-PstI homology 13ArxBeetle 
PrdPairedPax3/7Beetle 
PropPropPropBeetle 
PrrxPrrxPrrxBeetle 
Ptx1Ptx1PitxBeetle 
RepoReversed polarityRepoBeetle 
RxRetinal homeoboxRaxBeetle 
ShoxShoxShoxBeetle 
SvShavenPax2/5/8Beetle 
ToyTwin of EyelessPax4/6Beetle 
Unc4Unc4UncxBeetle 
VsxVsxVsxBeetle 
AlxAlxAlxAmphioxus 
Aprd1Aprd1AprdAAmphioxus 
Aprd2Aprd2AprdBAmphioxus 
Aprd3Aprd3AprdCAmphioxus 
Aprd4Aprd4AprdDAmphioxus 
Aprd5Aprd5AprdDAmphioxus 
Aprd6Aprd6AprdEAmphioxus 
ArxArxArxAmphioxus 
DmbxDmbxDmbxAmphioxus 
DrgxDrgxDrgxAmphioxus 
GscGscGscAmphioxus 
HopxHopxHopxAmphioxus 
IsxIsxIsxAmphioxus 
OtpOtpOtpAmphioxus 
OtxOtxOtxAmphioxus 
Pax2/5/8Pax2/5/8Pax2/5/8Amphioxus 
Pax3/7Pax3/7Pax3/7Amphioxus 
Pax6Pax6Pax4/6Amphioxus 
PhoxPhoxPhoxAmphioxus 
PitxPitxPitxAmphioxus 
PropPropPropAmphioxus 
PrrxPrrxPrrxAmphioxus 
RaxRaxRaxAmphioxus 
RepoRepoRepoAmphioxus 
ShoxShoxShoxAmphioxus 
UncxAUncxAUncxAmphioxus 
UncxBUncxBUncxAmphioxus 
UncxCUncxCUncxAmphioxus 
VsxVsxVsxAmphioxus